1. College of Horticulture，Agricultural University of Hebei/Key Laboratory for Vegetable Germplasm Enhancement and Utilization of Hebei/Collaborative Innovation Center of Vegetable Industry in Hebei; 2. Key laboratory of plant genetics and molecular breeding，Zhoukou Normal University/Henan Key Laboratory of Crop Molecular Breeding and Bioreactor; 3. College of Biological Engineering，Henan University of Technology

National Natural Science Foundation of China Youth Science Foundation Project（32002062）；The Key Scientific and Technological Project of Henan Province（202102110189，212102110423，212102110142）；Graduate Innovation Funding Project of Hebei Province（CXZZBS2019100，CXZZBS2019098）；The Training Plan for Young Backbone Teachers in Colleges and Universities of Henan Province（2021GGJS139）；Key Research Project of Higher Education Institutions in Henan Province（20A180034）