1Crop Research Institute, Shandong Academy of Agricultural Sciences / Key Laboratory of Wheat Biology and Genetic Improvement in Northern Huanghe-Huaihe River Valley / National Engineering Laboratory for Wheat and Maize, Jinan 250100; 2School of Life Science, Shandong Normal University, Jinan 250014
National Natural Science Foundation of China(32001542),Shandong Natural Science Foundation(ZR2020QC114), Open Project of State Key Laboratory for Wheat and Maize Engineering(2018LYZWS06),Open Project Fund of State Key Laboratory of Crop Biology(2018KF01)