Vegetable and Flower Research Institute of Chinese Academy of Agricultural Sciences/Key Laboratory of Horticultural Crop Biology and germplasm creation,Ministry of Agriculture,Beijing 100081,Vegetable and Flower Research Institute of Chinese Academy of Agricultural Sciences/Key Laboratory of Horticultural Crop Biology and germplasm creation,Ministry of Agriculture,Beijing 100081,Vegetable and Flower Research Institute of Chinese Academy of Agricultural Sciences/Key Laboratory of Horticultural Crop Biology and germplasm creation,Ministry of Agriculture,Beijing 100081,Vegetable and Flower Research Institute of Chinese Academy of Agricultural Sciences/Key Laboratory of Horticultural Crop Biology and germplasm creation,Ministry of Agriculture,Beijing 100081,Vegetable and Flower Research Institute of Chinese Academy of Agricultural Sciences/Key Laboratory of Horticultural Crop Biology and germplasm creation,Ministry of Agriculture,Beijing 100081,Vegetable and Flower Research Institute of Chinese Academy of Agricultural Sciences/Key Laboratory of Horticultural Crop Biology and germplasm creation,Ministry of Agriculture,Beijing 100081